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Study of interrelationships between organisms and their biotic and abiotic environment; includes the physical environment, population biology, community dynamics, biotic interactions and evolution. Usually offered Fall, Spring, Summer. (Cross-listed with BIOL 3340).

Prerequisite(s): Prerequisites are BIOL 1450 and BIOL 1750; junior-senior or Biology graduate student; or permission by instructor. Not open to non-degree graduate students.